This family is a lineage closely related to the Littorinidae (periwinkles) common in coastal habitat. They have adapted to terrestrial life and are thus sometimes called "land winkles".[1]
They are defined by a chalky operculum at the rear end of the body, the shape of their thick shell and their mouth and a characteristic spiral sculpture. The sexes are separate and can sometimes be recognised because the female shell is slightly larger than the male shell.
^Georgia in GBIF Archived 20 June 2015 at the Wayback Machine
^Guillainia in Bionames.org, accessed 12 december 2015.
^"Genus summary for Leonia". AnimalBase, accessed 28 march 2011.
^Lithidion in GBIF
^KENNETH C. EMBERTON Cryptic, genetically extremely divergent, polytypic, convergent, and polymorphic taxa in Madagascan Tropidophora (Gastropoda: Pomatiasidae) Biological Journal of the Linnean Society - Volume 55, Issue 3, pages 183–208, July 1995
^"Genus summary for Tudorella". AnimalBase, accessed 28 march 2011.
Pfenninger, M.; Véla, E.; Jesse, R.; Elejalde, M. A.; Liberto, F.; Magnin, F. D. R.; Martínez-Ortí, A. (2010). "Temporal speciation pattern in the western Mediterranean genus Tudorella P. Fischer, 1885 (Gastropoda, Pomatiidae) supports the Tyrrhenian vicariance hypothesis". Molecular Phylogenetics and Evolution. 54 (2): 427–436. doi:10.1016/j.ympev.2009.09.024. PMID 19772926.
Watters G. T. (2006). The Caribbean Land Snail Family Annulariidae: A revision of the higher taxa and a catalog of the species. 1-557, figs 1-10, map 1-56, Appendix A: 1-3. Backhuys Publishers, Leiden.